New Genus Voluceropecten and Family Voluceropectinidae of Late Permian Pectinids of the Boreal Superrealm

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Abstract

Based on the re-examination of the previously established species Aviculopecten volucer Lutkevich et Lobanova of the Late Permian (Late Wuchiapingian-Early Changhsingian) pectinids, which was considered a synonym of the Middle Permian Vnigripecten phosphaticus (Girty), the independence of both species is shown. Features of the peculiar shell morphology and sculpture of Aviculopecten volucer make it possible to describe the new genus Voluceropecten gen. nov. and the monospecific family Voluceropectinidae fam. nov. A corrected diagnosis of the genus Vnigripecten Muromzeva is given.

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About the authors

A. S. Biakov

North-East Interdisciplinary Scientific Research Institute n.a. N.A. Shilo, Far East Branch of the Russian Academy of Sciences

Author for correspondence.
Email: abiakov@mail.ru
Russian Federation, Magadan, 685000

References

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2. Fig. 1. Locations of representatives of the aviculopectinoid genera Voluceropecten gen. nov. (circles) and Vnigripecten Muromzeva (triangles) in the northeast of Russia: 1 - Southern Verkhoyanye, Kobyume River, left tributary of the Suntar River (upper reaches of the Indigirka River); 2 - Ayan-Yuryakh anticlinorium, Kolyma River below the mouth of the Detrin River; 3-10 - Omolon massif and its southeastern frame: 3 - Astronomicheskaya River, tributary of the Left Kedon River, 4 - Russkaya-Omolonskaya River, 5 - upper reaches of the Khudyana River, tributary of the Bolshaya Aulandzha River, 6 - left bank of the lower reaches of the Ledyanaya River, tributary of the river. Blue (interfluve of the Bolshaya and Malaya Aulandzha), 7 – Krutoy Creek, upper reaches of the Verkhniy Koargychan River, 8 – interfluve of the Irbychan–Korgychan River, 9 – Levyi and Pravyi Vodopadny Creeks, upper reaches of the Khivach River, basin of the Gizhiga River, 10 – Krutoy Creek, tributary of the Khivach River; 11 – Chistyi Creek (Fedorovsky), tributary of the Paren River. The dotted line shows the contour of the Omolon Massif.

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3. Fig. 2. Stratigraphic position of finds of representatives of the aviculopectinoid genera Voluceropecten gen. nov. and Vnigripecten Muromzeva in the North-East of Russia. Abbreviations: Lek. – Lekeersky, ISS – International Stratigraphic Chart, lower. h. – lower part, RSS – Regional stratigraphic scale.

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4. Fig. 3. Bivalves of the genera Voluceropecten gen. nov. and Vnigripecten from the Middle–Upper Permian deposits of northeastern Russia (Omolon Massif) and the USA (all images, unless specially noted, are given in natural size): a–l, f – Voluceropecten volucer (Lutkevich et Lobanova): a, b – holotype TsNIGR Museum No. 28/8448, core of a bivalve specimen with separated valves and partially preserved shell: a – from the side of the right valve, b – from the side of the left valve, Levyi Vodopadny Creek, Khivach River basin, c – specimen TsNIGR Museum No. 30/8448, imprint of the right valve with clearly visible spines on the hinge edge, same location and age; g–e – specimen SVKNII #114/02–06.99, incomplete core of a bivalve specimen with a partially preserved shell: g – from the umbo, d – from the right valve (arrows indicate: gs – gill suspensoria, za – posterior adductor, ml – mantle line, pm – pedal muscles-elevators, pr – pedal retractor), e – from the left valve, Pravyi Vodopadny Creek, exp. 9, layer 3, author’s collection, 1983; g – specimen SVKNII #113/02–06.99, core of the left valve with a partially preserved shell, same location; h – specimen. SVKNII #46/02–06.99, core of the right valve with a partially preserved shell, same location as in fig. a, outcrop 8, layer 4, author’s collection, 1983; i – specimen SVKNII #112/02–06.99, core of the left valve with a partially preserved shell, upper reaches of the Khudyana River, tributary of the Bolshaya Aulandzha River, so-called 1357, collection of L.D. Shkolny, 1986; k – specimen SVKNII #115/02–06.99, incomplete core of the right valve, Pravyi Vodopadny Creek, outcrop 3, layer 2, author’s collection, 1999, see designations for fig. d; l – specimen. SVKNII #44/02–06.99, right valve, Russkaya-Omolonskaya River, outcrop 32, member 22, author's collection, 1983; f – specimen KFU #551/47, incomplete imprint of the right valve with a partially preserved shell, Kobyume River, left tributary of the Suntar River (upper reaches of the Indigirka River), the structure of the hinge area with the orthoclinal position of the ligamentous pit is visible; a–c, z – upper part of the Khivach Formation, Upper Permian, lower part of the Changsing Stage; Kolyma Superhorizon, Khivach Horizon, Intomodesma costatum bivalves zone, upper half of the I. evenicum subzone; g–g, k, l – upper Khivach Formation, upper Wuchapingian Stage; I. costatum bivalve zone, lower half of the I. evenicum Subzone, i – upper Armandzhin Formation, upper Wuchapingian – lower Changhsingian Stage; I. costatum bivalve zone, f – upper Privolninskaya (Imtachan) Formation, same age; m–u – Vnigripecten phosphaticus (Girty): m – specimen USNM #140473, left valve, ×1.3, Mid Peak Member, Phosphoria Formation, middle Permian, Coal Canyon, near Cokeville, Wyoming; h – holotype USNM #1737, cast of left valve, ×2, phosphate beds of the Phosphoria Formation, same age, Thomas Fork, Wyoming; o – specimen USNM #140475, right valve, ×1.3, same locality and age as fig. m (all images from: Ciriacks, 1963); p – specimen SVKNI #116/02–06.99, core of right valve, Irbychan – Korgychan interfluve, middle Permian, Roadian Stage, Aphanaia dilatata Bivalvian Zone, lower part of the Avlanda Formation, so-called 5240, collection of V.A. Kovalchuk, 1988; p – specimen SVKNII #117/02–06.99, core of the left valve, the same location and age; c – specimen SVKNII #118/02–06.99, core of the left valve with sections of preserved shell, Krutoy Creek, upper reaches of the Verkhniy Koargychan River, lower parts of the Omolon Formation, the same age, so-called 37-10, collected by N.K. Karavaeva, 1979; t – specimen SVKNII #119/02–06.99, core of the right valve with a partially preserved shell, Krutoy Creek, tributary of the Khivach River, upper parts of the Dzhigdalinskaya Formation, so-called 06-10, the same age, collected by L.D. Shkolny, 1983; y – specimen SVKNII No. 120/02–06.99, incomplete core of the right valve, left bank of the lower reaches of the Ledyanaya River, tributary of the Sinyaya River (interfluve of the Bolshaya and Malaya Aulandzha), lower part of the Avlandinskaya Formation, the same age, collected by N.K. Karavaeva, 1986.

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